!"#$!%&/!-0($&'( )*+,&%!*"(-. !"#$% $& '#$(!#)*' +,-+. !"#$%&"'(' '$ )( )(*+!( )( )(*+!(, -. '$ !/#"$01%$ '$ 2..3 ! "#$%&' () *# +,-./&(#(! !" #$%&!'("( () m )*+)#')* N1, . . . , Nm ,$-).!"(" +$. )/ *'*0 1)%"2 dNi = Fi(N1, . . . , Nm) dt 1 ≤ i ≤ m, 3 #$!4,&."#'5! () )6&'/'-.'$ N ∗ = (N1∗, . . . , Nm∗ )2 ∗)=0 Fi(N1∗, . . . , Nm 1 ≤ i ≤ m. 7!8/'*'* () /" )*1"-'/'("(9+).1&.-"#'$!)* ()/ )6&'0 /'-.'$2 Ni = xi + Ni∗ dxi ∗ +x ) = Fi(x1 + N1∗, . . . , Nm m dt m X ∂Fi (N ∗)xj + t.o.s. = j=1 ∂xj = :#&"#'5! /'!)"/';"("2 m X dxi aij xj = dt j=1 m X aij xj + t.o.s. j=1 dx = Ax. dt • A . ! "!#$%& '( /!/( '( *0 ! )*"+,%'!'- aij 1 aji2 m X dxi aisxs = · · · + aij xj + · · · = dt s=1 - m X dxj = ajr xr = · · · + ajixi + · · · dt r=1 aij = (3()#* (0/()%( j 0*4$( 7 8 7 77 78 8 87 88 − ↑ σ(aji) −+ −0 • ++ ! 5!$%!)%6, (0/()%( − +− 0− −− ← σ(aij ) "+#+! %0"*1 0%"4%*0%01 )**/($!)%6,- • −− )*"/(#%)%6,- • +− /$(0!9'(/$('!'*$- i- ∂F • ∂xj ≡ 0 i $"%&" j / ∂F • ∂xj ≥ 0 i ! "#$"%&" $!-! xi '( &')*+" #(,-" ! "#. i 6= j 0 i, j !-,&1-!-&(#2 #&#1"3!# i 6= j 0 i, j !-,&1-!-&(#2 #&#1"3!# %(($"-!1&4(#/ ∂F • ∂xj ≤ 0 i $!-! %(3$"1&1&4(#/ ! "#$%&'($ )*'+&%#%#,*$! # " dN1 N1 + b12N2 = r N 1 − 1 1 dt K1 # " N2 + b21N1 dN2 = r N 1 − 2 2 dt K2 • Ki !"! #$!$ $% &'"'()% "!(! *! %&"% #% i +# N1, N2 $'& %&"% #%& $% -%(./0'('&, *! %&"% #% 1 2!3'( 45% *! 6 %7)'7 %& K1 < K28 !"! #$!$ $% #7)(5&#97 $% *! %&"% #% 6 &'.(% *! 1, "'( :57#$!$ $%; #7$#0#$5'8 • b12 !"! #$!$ $% #7)(5&#97 $% *! %&"% #% 1 &'.(% *! 6, "'( :57#$!$ $%; #7$#0#$5'8 • b21 <$#2%7&#'7!*#=! #97> Ni , ui = Ki τ = r1 t r2 ρ= r1 du1 = u1(1 − u1 − a12u2) dτ du 2 = ρu2(1 − u2 − a21u1) dτ b N • a12 = 12 2 K1 , !"#$% &! !'"( )' *( '%+', ' - %./#' *( 01 +.# $! )() )' #',$#%.% ) %+.! /*'% +(#( *( '%+', ' 02 b N • a21 = 21 1 K2 !"#$% &! !'"( )' *( '%+', ' 0 %./#' *( -1 +.# $! )() )' #',$#%.% ) %+.! /*'% +(#( *( '%+', ' -2 !"! #$%&'&()&*+ • (u∗1, u∗2) = (0, 0) !"#$%#&$ ' ()* '+* *, %# *• (u∗1, u∗2) = (1, 0) !"#$%#&$ ' () *, %# .• (u∗1, u∗2) = (0, 1) !"#$%#&$ ' () *, %# /• 0+ !#*" $%#) (u∗1, u∗2) = Ã 1 − a12 1 − a21 , 1 − a12a21 1 − a12a21 (+ 12 3 12# 3 4 a12 > 1 & a21 > 1 a12 < 1 & a21 < 1. & ! !"#$!%&'!() *+ ,-" .&)#-" *+ +/&!,!%$!-0 • (0, 0) ! "# !$%&' (#)*)+, • (1, 0) ! !$%&' !" '% -%.%-"*%* * "#$/0!"1# * '% !. -" 2 !)&/ '% 3 !0. /% '% -%.%-"*%* * !).)/$ * 4!$%, (1, 0) = !$%&' !" a21 > 1 "# !$%&' !" a 21 < 1 . • (0, 1) ! !$%&' !" '% -%.%-"*%* * "#$/0!"1# * '% !. -" 3 !)&/ '% 2 ! !0. /")/ % '% -%5 .%-"*%* * !).)/$ * '% !. -" 2, (0, 1) = !$%&' !" a12 > 1 "# !$%&' !" a 12 < 1 . ! !)!$ #"&' ( !5 $%&' + !" '%! !. -" ! #) %8)$%# '%! -%.%-"*%* ! * !).)/$ * '% !. -" ).)# #$ aij < 1, 9 5 !0'$% "# !$%&' !" %:&%! !. -" ! * !&)/*%# '%! / !. -$";%! -%.%-"*%* ! * !).)/$ , • 6% -) 7"!$ #-"% (u∗1, u∗2) !"#$%&%'#' ( )*+,*-"#+%./"* #!%/"0"%)*1 2-%/)%,%* '. .3)&4!%0/ )*+,."%"%5#6 )#!*! 7$89 7)89 7'81 ? ? :*.3%!"./)%# # ,.!#- '. &# )*+,."%)%0/1 ! "#$#%&'()* * (')+'*('(! " # N1 N2 dN1 = r 1 N1 1 − +b12 dt K1 K1 " # N2 N1 dN2 = r 2 N2 1 − +b12 dt K2 K2 • Ki !"! #$!$ $% &'"'()% "!(! *! %&"% #% i+ ,$#-%.&#'.!*#/! #0.1 Ni , ui = Ki τ = r1 t r2 ρ= r1 du1 = u1(1 − u1 + a12u2) dτ du2 = ρu2(1 − u2 + a21u1) dτ , • 234#*#5(#'& #.%&)!5*%& (0, 0)6 (1, 0)6 (0, 1)+ • !"#$#%&#' () *')+#,-).*#/0 1 + a12 1 + a21 ∗ ∗ , , (u1, u2) = δ δ ('.() δ = 1 − a12a211 2,#.-3-#*/4).-) ,# δ > 01 µ ¶ ),-/%$) 5"/.(' δ > 0 .' 6/7 *')+#,-).*#/ 7 $/, (', ),8)*#), )+8$'-/. ). -#)48' 9.#-'1 • ! "#$%#&'($)*+'# #,-).('#&)#/ ,#.0-)#+*, %*&)12)-(,! !" #$%&'()! *+,()-('" N ∗(t + T ) = N ∗(t) -+ &! #(#.+/" dNi = fi(N1, . . . , Nm) dt t∈R i = 1, . . . , m #+ -('+ 0$,1(."%/+!.+ "#(!.).('"/+!.+2 +#."1%+3 #( +# +#."1%+ 4 #( "-+/5# ∃η > 0 ."% 6&+ *"," .$-$ |N0 − N ∗(0)| < η +7(#.+ τ > 0 '$!8 lı́m |N (t) − N ∗(t)| = 0. t→∞ 9:;: <$%&'($!+# *+,()-('"# +! *%"!$: 3*#&*$( 4 =>$(!'",?@A+!-(7$!B! !" N (t) #$%&' i = f (N , N ). ()*+ ,! %" !(&"()*+ C 1 -%"+" dN 1 2 i dt i = 1, 2/ ) N (t) ∈ K -"0" t ≥ 0. ,$+,! K !# &+ ($1-"(2$ #)+ -&+2$# ,! !3&)%)40)$. !+' 2$+(!# !5)#2! &+" #$%&()*+ -!0)*,)(" N ∗(t) !+ K 2"% 3&!6 |N (t) − N ∗(t)| → 0 t → o∞. • !"#$ #%&!'( )*!+','!")&(+'( (*')-#(. !"! #$%&'(%&)*+$ ,-./&0($ ( 1* +21&'&3-&)! !"#!$% & 45),67' !" N ∗ #$ !%#&'&()&* +! '" ,"-&'&" +! !.#".&*$!/ C k 0 k ≥ 20 -1+&-!$/&*$"'!/ dNi #3*$4"-*/ dt = fi(λ, N1 , . . . , Nm)0 λ ∈ R2 %#!5 67 ∂f µ(λ) ± iω(λ) ∈ σ(A(λ))0 A(λ) = ∂N (λ, N ∗)2 87 µ(0) = 00 ω(0) > 00 σ(A(0)) ∩ {<z = 0} = ±ω(0)i2 97 dµ dλ (0) 6= 02 :$;*$.!/ !<&/;!$ ,#$.&*$!/ λ(s)0 T (s)0N (t, s) 3!)&=+&." +! 3!)&*+* T (s) > =)(&;" Γ(s)0 |s| < ε0 > #$ !$;*)$* U ⊂ Rm +! N ∗ ;"'!/ %#!5 &7 N (t, s) /*'#.&=$ 3!)&=+&." 2π 2 λ = λ(s) .*$ T (0) = ω(0) .*))!/3*$+&!$;! " &&7 Γ(s) !/ '" ?$&." =)(&;" +! 3!)&*+* T (s) .*$1 ;!$&+" !$ U 2 ! "#$%&'($ )*&$(+,&-*&.(./* /$0#1(%/*#/$! µ ¶ N dN = rN 1 − − P N R(N ) dt K µ ¶ hP dP = kP 1 − dt N R(N ) !"#$!"%& '$()*+(&, -!, -!# !-&-+ & ,& .& *&)*/( !( !, (01! + -! # !"&"2 3#)*+(!" #& & N R(N )4 a) = AN 4 b) = AN/(N + B)4 c) = AN 2/(N 2 + B 2)4 d) = a[1 − e−aN ]2 !"# $%&'('% )* &#)*(#+ %(!*",%!'-#+. dN = rN N 1− K µ ¶ − P N R(N ) dt dP = kP (−d + eN R(N )) dt /% #01'2, 34 ¶ µ dN AN N −P = rN 1 − dt K N +B µ ¶ hP dP = kP 1 − . dt N 5)'&*,+'#,%('6%1'2,. N u= , K a= A , rh h v = P, K k b= , r τ = rt d= B . K du v = u (1 − u) − a dt v+d µ ¶ u dv = bv 1 − . dt v !"#$#%&#' (u∗, v∗)( u∗2 + (a + d − 1)u∗ − d = 0. u∗ = v ∗, u∗ = (1 − a − d) + )*+%#$#,+,( u A= ,-* ∗ · au∗ (u∗ +d)2 q (1 − a − d)2 + 4d 2 ¸ −au∗ u∗ +d −1 b " . −b # . ad ∗ > 0. A= + 1 bu (u∗ + d)2 (u∗, v ∗) = -)*+%$- )# .&+/+A < 0 #0-)*+%$- )# .&+/+A > 0.