Metazoos LE 32-3 Single cell Stalk LE 32-4 Somatic cells Digestive cavity Reproductive cells Colonial protist, and aggregate of identical cells Hollow sphere of unspecialized cells (shown in cross section) Beginning of cell specialization Infolding Gastrula-like “protoanimal” Filos animales s/estructura corporal I. Parazoa: poríferos (esponjas) (sin verdaderos tejidos) II.Eumetazoa (con auténticos tejidos) A. Animales con simetría radial (diblásticos): cnidarios B. Animales con simetría bilateral (triblásticos): 1. Acelomados: platelmintos 2. Blastocelomados:nemátodos 3. Celomados a. Protostomados: moluscos, anélidos y artrópodos b. Deuterostomados: equinodermos y cordados Simetría Radial Simetría Bilateral LE 32-8 Coelom Body covering (from ectoderm) Digestive tract (from endoderm) Tissue layer lining coelom and suspending internal organs (from mesoderm) Coelomate. Coelomates such as annelids have a true coelom, a body cavity completely lined by tissue derived from mesoderm. Body covering (from ectoderm) Pseudocoelom Muscle layer (from mesoderm) Digestive tract (from endoderm) Pseudocoelomate. Pseudocoelomates such as nematodes have a body cavity only partially lined by tissue derived from mesoderm. Body covering (from ectoderm) Tissuefilled region (from mesoderm) Wall of digestive cavity (from endoderm) Acoelomate. Acoelomates such as flatworms lack a body cavity between the digestive cavity and outer body wall. LE 32-9a Protostome development (examples: molluscs, annnelids, arthropods) Eight-cell stage Spiral and determinate Deuterostome development (examples: echinoderms, chordates) Eight-cell stage Radial and indeterminate Cleavage LE 32-9b Protostome development (examples: molluscs, annnelids, arthropods) Deuterostome development (examples: echinoderms, chordates) Coelom formation Coelom Archenteron Coelom Mesoderm Blastopore Schizocoelous: solid masses of mesoderm split and form coelom Blastopore Mesoderm Enterocoelous: folds of archenteron form coelom LE 32-9c Protostome development (examples: molluscs, annnelids, arthropods) Deuterostome development (examples: echinoderms, chordates) Mouth Anus Digestive tube Mouth Mouth develops from blastopore Anus Anus develops from blastopore Fate of the blastopore Protostomos Deuterostomos Blastoporo origina la boca Blastoporo origina el ano. Segmentación espiral Segmentación radial Formación celoma: esquizocelia Formación celoma: enterocelia Cadena nerviosa ventral – Gastroneuralia Cadena nerviosa dorsal Notoneuralia Desarrollo embrionario temprano •Fertilización y formación del cigoto • Segmentación y formación de la blástula •Formación de la gástrula Clivaje reductivo Patrones de segmentación De acuerdo al tipo de huevo: Holoblástica o completa (isolecitos, mesolecitos) Meroblástica o parcial (telolecitos, centrolecitos) Gastrulación fibras Movimientos morfogenéticos Células mesenquimáticas Gastrulación •se establecen los ejes corporales •movimientos celulares •aumenta el número de capas celulares •se forma el arquenterón (tubo digestivo) •aparecen signos visibles de diferenciación •aparecen las tres capas de tejido embrionario: a. Endodermo: interna b. Mesodermo: media c. Ectodermo: externa Larva pluteus LE 32-10 “Radiata” Deuterostomia Protostomia Bilateria Eumetazoa Metazoa Ancestral colonial flagellate LE 32-11 “Radiata” Deuterostomia Lophotrochozoa Bilateria Eumetazoa Metazoa Ancestral colonial flagellate Ecdysozoa Figure 32-12 LE 32-13 Apical tuft of cilia Mouth Anus An ectoproct, a lophophorate Structure of trochophore larva