Fermentación maloláctica

!"#
$"
#" &
'( #!"!
*
%
!
" & *
!
!"#
') '"
!
/!
$
%&
" &
#*
! '
)
)
12
" !
'
*
#
#$
(
( +
#
#(
*
'- *
(
" & -'
!
/!
0
'0 3"$4#
.
*
0
*
)
5
# (
0
.
!
)
8
!
(
*
9
1:2 / !
!) /
;
(
(
" &
!
0
* '
!*
(
#
<
=( / !> < ? -=( / !> <5# !
*
0
3@ A4#
C&
' %
+
,
; (
* -
-(
0
(
)
*
#- . ' - *
0
!
*
,
(
*
/!
!
'
*
/!
)
162#' 7
!
)
(
#*
(
)
!*
!
!*
0 4' '
#
/
"$ 3
!
(
!*
0
!
(
* & ! #0 3B4
-
!*
)
-
!)
!
" & '
*
*
*
!*
0
! /!
0
'0
- * &%"# '"
&
(
*
*
.
( /
(
#
-0 ' ! 7 !
!
6DDE - 6D D
<
*
(
;
*0
DD& %
(#
'
*
@
!*
66&E
8*
/!
F
; (
3< ?C@ A4 -
=( / ! < ?> 3< ?C B 4 -
)
* ;
(
=( / ! <5> @ A - <5C B 4 & !*
%
/
# /
#
/
-+
'(
)
66&E ! 7
!
/!
0 /
66G
/!
'0
-!
)
#
- !
" ! & !(# /% ' #
(# *-+ *
*
!
'
0 3<5C
0 EEH
(
/
#
&0' ' !(# ') ! & 1#2
') 2 &"&0' '
(!
(
)
( ! 6DDE
(
(
* (
<
( !
6D D
(# *
3?4 & / '
(
(# +
3$4
! !
3?4I
)
& / '
0
0
! !
0
(
3$4-
)
0
0
0
(
)
6
$ ! !
!*
- - 2$&
B
-
/!
0
(
!
(
!
" &/
0
6DDE
') ! & % ' #
)
$ C
# !
)
0
0
" &
)
* )!
0
*
(
@A- B
*J -)
!)
/ (
!
B *
!
*
%& *F
)!
!
&0' '
!
!*
*
!*
0
!
/ C !
%& * @
(
!
(
/
*
( ) #)
)
* )!
/ C
! #
(
7
/
!
*
(
3@ A4-!
!*
0
" &
/
'0
/!
& (
#
@
:#
MN
@
D#
6:
:#
E:
D#
@
:#
H
D#
::
D#
D
D#M
:#
D#
DK
:#
DN
D#
DK
:#
H:
D#
D
K#
66
D#
D:
K#
6N
D#
DD
K#
6:
D#
D:
D#
:D
D#
D:
D#
:
D#
D
D#
K
D#
DD
D#
H
D#
D
D#
D#
D6
D#
D#
D
!)
)
$
J
6
!
% !
6
6
(
"
6
!
O6 ! &
DDH
.
6
/
D#
DD
D#H
D#
DD
D#
D#
DD
#N
D#
D
#
H
D#
DD
D#
:
D#
DD
D#
DK
D#
DK
D#
:6
D#
DD
D#
DD
D#
D
D#
DD
D#
D
D#
DD
D#
D6
D#
DD
D#
6E
D#
K:
D#
DM
#
DE
D#
HK
D#
KE
D#
DK
K#
K:
D#M
K#
6
D#
DN
K#
KE
D#
D
K#
D#
6E
D#
HE
D#
D:
D#
HD
D#
DD
D#
HE
D#H
D#
H6
D#
DK
D#
D
/
D#
D
D#
DK
(
D#
DD
PQ
#
6H
D#
DD
D#
DM
D#
D
D#
D6
#
DE
!
@ A*
(
*
B
:#
D:
$
#
<5
@A
K#
6:
$
#
(
'
*J
( )
.
!
$
$
F
!
0 /
!
-/
!
B
D#
'
0 3B4
@
:#6
!
-)
*
-
< ?
' 3L((4
0
!
!
(
'
- !
< ? -<5
@A
$
(
D
-3
0& ! &" $ #02 #" 14 '" 5(42 '" ! &"
"
& (
/
!
& * )!
'0 #
#*J#
( ) #)
!) #)
)
.
!7
/
)
1H2 &
!
0
!7
* 1K2
&
(
(
#
6N
D#
DH
D#
D
D#
DK
3# # 4
*
@
C5 I! CR
D#
D
/ (
:
- 6 "2$" ')
"&0 &! &"
&
!*
( )
!*
0
* 8
0
0 *
/
"
(
C
0
/
*
!*
HDD!& (
! <
8
*
*
*
%& +
8*
!*
(
5
'
(
/
(
#
#
' #)
- !*
(
! '
#
'
# *
- !*
7
(
*
0
*
//
#
#
-/
( )
&
'
B
/
(
)
!
@ A*
! - 8* 0
!
* <5
(
*
( *
!*
%& +
<5
3B4
(
B *
!
$
:CBJC6C
$ '
?
$ '
6
%
'
$ '
7
6
" /
S
$
T
@
6
5
*
6
% *
&
!*
7
6
(
<
6
"
( )
! O6
O /
*
@
C5 I! CR
( )
-*
!
(
#
B @
! -
(
'
!
!
#
@ '
7
#
(
!*
!)
#/
* &
" &
(
(
;# !
-)
-/
#(
*
!
(
*
3@ A4- !
!
(
< ?0
7
@
K#
EM
HN#6
K6#
N
E#
NH
6#
N6
:NM#M
6K#K
:#
:H
HHD#
D:
H#
D
H6#
HN
E#
HN
6#K
:D#
:H
D#
6:
D#
EH
66#6
#
H#D
N#
KM
E#
EK
(
!
"
*
-
( )
" & '
'
/
* < ?
7
$
@
8
C
D#
6:
D#K
N#
HN
D#
6D
D#
DM
:#E
N#
H:
D#:
6M#
D:
D#
D
#
MK
D#
6
D#
KM
#
:K
D#
D
D#
DK
D#
M
D#6
D#
DM
#H
D#
HM
3# # 4
@
:#H
K #
ME
#
:
D#
K
6#
K :#
MD
66#
MK
K#
::
6ND#H
M#E
:6#
H:
D#E
6#
6H
6E#
KN
D#
:K
#
6
:6#
NN
#
H6
M#
ME
MH#
:H
E#
6K
/
D#
H
D#
K
#
:E
D#
M
D#
6N
:#
K
#M
D#
:6
D#
:6
#
KK
D#
DK
D#
D#
DM
#M
D#
D:
D#
D:
#6
D#
6:
#
KK
:#D
D#
H6
/
@
6#
HM
D#K
K#
:H
HK#
MD 6#
6:
KN#
EK
K#
6E
D#
:N M:#
KM
H#
:H
D#
D
#D
D#
MH
D#
DE
#6
:D #
HH
6#
K
: K#
M
K #
:E
D#
KM 6E#
HH
K#
DE
D#N
K#
HE
H :#
HH
6D#
H #
ED
#
6
D#N
6H#
MN
KM#
D6
#
N
6E#
EM
E#
H6
D#
KD
#
ED
#
EN
D#
D
6#H
6N#
M6
#
MD
6H#
D
D#
:6
D#
D
D#
K:
#
DM
D#
:
#
D
6:#
ME
D#
HH
:D#K
#
KN
D#
6E
#
DM
#
:
D#N
6H#
NM
M:#
K
6#
M
#
KH
E#
:E
#H
M#
N6
(
(
PQ DDH
@
#
::
K#
MM
6#K
D#H
D#H
N#
:H
:#
H:
D#
DE
N#
N
#
#M
#
:E
D#
D#
KD
D#
6
D#
K
6#
H
D#
D:
#M
K#
:6
D#K
.
K
-8
0&
)
! !
"# &
@
*(
0
/'
/
0
* ! -
(
0*
*
/!
*
6
(
N@
*
!
#/
(
#(
0
-)
!"# $
!" %
(# - $ )
" & '
3
@
*(
!
9
:
(
3@ A4-!
0 3B4
*
9,
.
(
< ?
3 * (%&' '"
#
#7
#"#
! # #@
#@ #
*J
#
& 36D 64 1&( ' "1 "'(& " 2 "1
( "'; ;" "(
&' ! 2 &"&' '
% ' #(2 " "& & !
"#/ $#"1& "1 2$# &&" !
#&" <
9
/"
-# H #6KHC 6HK
5(
/?
3+
#
#
& "'(& "
?
# '$ ')
#
! # #7
'" "'(& "
3 (
*
" # "
#"#@
%#
# '<
*
04
#@ #
1#2
*J
" 9
#
# '" &
#
! # #J
#9 # 7
#"#@
#
@#
*J
#
&
#2
') 2 &"&0' '
"2$ # ') ! & "'(&')
'( ' &/ & '" "'(&') & !(# % ' #
0
FU
5
(
0
3
4#97
"
# : ;
6D
H
6
:
1 2?
#%O?
<
#$ EEH
M
#$ O
2
#9 O%
% " = !(
162 R
# @$
J ' ' != (
ENE B* !
-#
/ $
!
# &O
> %"& 2
'
!
&
1:2 9
# O
#$ O
#
"'(& " < ; /
! % ' # " 1#2
'""&'&2
'; #!" /
B
#9 OR
"?' >
I
V
C
M MCMM CK6C
J
'
#
'
#
6DD
11' "1 2(& "(
" @
' ! @ /<
$ #2 #
66 C66M
1K2 0! C$
#@ OJ !
# V
C ! # 6DDM 2(&
"1% "
' 2
9 2 " '!
! 22" (2 "
2 " "
! % # 2$&
HH DNW :
1H2B < 36DDE4
!
B <#
-#
"( A
!#
&/
<