Origen y Evolución de los Dinosaurios

Anuncio
Centro de Estudios Paleontológicos de Chile
!" # $
#
%&'&
! "#$%
"
(
)
*
(
*
+#
.
,
'45%6#
8
-
/ ,
#
1
#,
7
29 )
#: )
1
+,
,
0
';<
,
)
0
23
6
-
#
INFERIOR
MEDIO
TRIASICO
SUPERIOR
"
)
2"
,
# %=> %%4
INDUANO
251
249
OLENEKIANO
249
245
ANISIANO
245
237
LADINIANO
237
228
CARNIANO
228
216
NORIANO
216
203
RAETIANO
203
199
,
9 )
$
2
)
6#
,
9 )
8
9 )
/
! (
2
,
# %5< %=>
/
6 ?@
# %%4 %';
8
$
6#
,
# %&'& @
A
5;5BC< C4D
&
' ( )
*
,
9 )
#
8
(
#
7
2
-
(
#
%5&
*
,
/
- #
(
(
1
#
#
0 /
0#
#
6
,
# )
(
,
) /
7
#
- #
(
+
?
#
INFERIOR
MEDIO
TRIASICO
SUPERIOR
8
#
# %&&4D
INDUANO
251
249
OLENEKIANO
249
245
ANISIANO
245
237
LADINIANO
237
228
CARNIANO
228
216
NORIANO
216
203
RAETIANO
203
199
' +,
(
(
,
(
0
7
'
B
/
%
#
- )+
.
0
(
' E
% E
)
(
/
,
+ (
)
(
( .
F
F
- )+
"
/
#
/
,
/
, (
/ 0
0,
#
"
)
7
#
0
1
(
1
G 0
#" /
# ) ';# 'CC=D
3
/ #
(
)
,
1
/
?
7
(
#
0
(
(
#
#,
)
/
$ H !$
@
#0
/
)
)
#7
3
,
! I 3 @!9 !$
! 2$ "I# '4446
1
/
7
.
/
$
#
(
03
,
2$
/
0# '4446
(
$
,
0
/ 2
/
,
/
,
#
2
3
(
6,
(
$
G
6#
/
/ 6
G
/
,
2
G
3
G
(
,
#
)
$
(
JJ ,
)
0
/
/
1
#0 ,
#
(
7
3
/
#$
2
/
( .
(
/
,
,
/
#
(
1
7
"
3
/
/
0$
(
0
(
2
/
/
60
6
0
0
$
#
( .
1
,
,
#
(
/
/
(
#
(
0
JJ#
!
.
1
# 1
/ #
-
0( /
- )+
-
(
,
(
(
/
' !
#
, .
0, .
% !
/
1
= !
5 !
1
/
/
/
G
/
/
#,
( 60
2
(
(
/
#
#
1
2
(
6,
'0%
,
=
)
"
#
5#
,
00
9 )
$
8
9
,
K LL
0M
03
$
,
2
(
7
)
K LL
2'C>56
K
60,
G
(
2'C><6
2'C5<6#
" $ $!@ 383 N! $ A! @ !
$ 3 $ H !$ ! I 3 @!9 !$ ! !@ ! @ OH
8K3$ "!@ : $ 3$ @ H@ @9
$ 3 38H@ 8 $
! @9 OH 3@ H "OH!
39 3
3$ H !3 "3 H " 3@$9!9HI
!@3$ H ! 383 H@ M H 3 @ 9H " 3
83@3N!" 9! 3
,
+ /0 ;;
*
$
0#
< , (+
0
! "35$
/
(
/
,
0
,
7
,
# K LL
0
,
0
,
/
"
(
#
1
0
)
(
(
2 )(
9
+
! "33$2
/
B
N
1.2
3
,
,
(
0M
)
#0,
)
4 %"5! 67 2 89:
6
) + 0 ( -
"
.
(
(
)
,
7
(.
1
/
,
7
)
(
,
)
#
) 0 = . /
(
.
)
(
.
(
#,
G
1
#
,
#
0
1
(
#
,
(
.0 (
)
+
/
+
(
-
o
+
,+,
( /
"
:
.(
(
0 2M
# 'C4;P K
Q
'CC5P @ / 'CC; P K
# 'CCC6#
0
# 'CCC # 'CC>P $
L# 'C44P $
Q
# 'CC=# 'CC5P
(
1
(
7 #
0
/
/
(
G
/
/
0
#
,
K LL
0M
- #
,
(
/
'C>5
()
G
#
7
(
/
0
/
#
(
#
(
0
,
/
0
(
#
)
)
(
RH
-
.#
'C><# : . K
#
8
, -
,
0
#
@
P
.
(
#0
2S6 1
/
G
R 1.2
(
)
ó
6
>6
6
@
> >í?
0 AA "3#B 2
2S6
ó
#
! "33$2
4 %"5! 67 2 89:
( ó
/
(
/ í
(
0
ó
(
á
$
%=C
"
9
"
0 !! 9
# 2%&&<6 :
($
!
$
'C #%'C–
.
" N
N
!
"
1
!
(
+ !.
,
(( ! "3$#
7
#!
N
)C ,
(
N
"
/ )
0"
"
#
!
,
9 )
7
(
0
1
N
#
!
(
0
( /
0
0 N
8
!
N
#N
9 )
"
#
1
"
0
0
$
#
#
)C
"
) - ! "#B3
/
,
7
)
,
&#> %5<#& T '#<
/
/
/
9 )
/ (
- 60,
23
#
L
# %5C#> T
8
(
9 )
2K
# 'C'=6#
, //
2%5> U %5<
60,
#
,
/
. ( ,
)
(
+ +
"
-
)
(
3
)
,
L
/
29 )
+
+ (
6
(
)C
"
)
0
G
0
0
#
#
P
/
0
#
1
7
#(
0
G
)
P
/
0
/ # ,
)
)
0
#
,
0
# (.
1
)
(
,
P
+C
"
,
! "3#B
0
/
0
8 0
#
(
/
,
/
/
/
/
)
)
/
(
,
+C
#
,
3
"
2
( 1
( 1
0
/
6# ,
)
0
#
/
R$R
(
/
0
,
(
(
/
(
#
,
0
3
2V
,
H
-
9 )
/
9
é
,
/
,
,
1
#,
(
/
1
#,
0
(
í(
/
0
# 'C&>6 #
#
#
ó
í
#
"
1
7
(
#
0
/
> ?
"
"
/
> 6
2
-C
6#
( ,
#
2
(
(
9 )
0
$
2@
#0
@
@
@
6
)
#
/ 8.1 #
K
#0
+
! "3#$
#
0
6
9 )
#
0
8
2"
# @ / 8.1
0
0/
G
.
(
6
09 1 9
9 )
"
,
0
#
.
$
0
2
9 1
0
#
6
#0
.
/
> ! "35"
-
2
# 'C>'6#
1
0
7
,
/.
2'C>%6
(.
#
#
K
0(
2'C456
/
$
(
/
,
(
#
7
0
.
/.
#
7
#@
,
(
/
#
0
#
G
,
)
,
)
/.
,
1
)
(
,
/
0
#
0
# %&&>6#
"
(
2K
7
.
9 )
9
2
#'C4;6#
2K
# 'C4<6 0
2@ / # 'CC%6 ,
# 'C><6 0 (
.
2!
(
@
# %&&> @
. #
N
)
/
/
"
(
)
(
0 (
/
(
2 A" 5;'C# H " &;6
/ # K#
/
#
2 A" 5;'C6
((
(
(
H " &;
&;6 /
/ 2 A" 5;'C6
/ #
#
( (
/
#
0
/
(
N#
2 A" 5;'C6 M#
(
#
/
:#
H " &; W#
(
#
/
(
2H "
!A
(
A" 5;'C
H " &;
/
%P
5 B(
# B
P B
P '# %B
'# %P ! !AB
! !AP ( B (
P B
P B
P ! AB
! AP
B
P B
P ' %B
'
B
P B
$
,
'
/
(
/
P
A" 5C';
&; @
H "
.
/
'
! "33%
(
2
# 'C>'6#
/
#
)
/
0
#
,
0 ,
#
,
? >
"
/
7
,
.
0
0
#
"
G
/ #
#
(
> ! "35"
#
,
K
#/
/
,
7
#
)
# 'C>%6#
"
#
,
0
(
2'C><6#
7
#
G
"
7 #
0
/
/
0,
7
,
/
)
,
-
)
1
/ #
/
0
<&X
K
/
2'C>%6
#
2
.
0
,
'C;5
,
G
)
#
<(
0
/
#
(
,
,
(
,
#
B
'
0
%
0
0
(
(
(
=
5
0
(
0
#
0
)
)
>
"35% D
E
! "33$
)
$
H /
#7
#
0K
#
"
,
/
H /
/
/ #
#
/
/
7
"
#(
(
2
.
/
,
7
29
$
'' 'C $
#
#
K
6B
! (
#
/
,
$
#
/
(
(
7
#0
'C>% Y $
2
"
(
# 8 : 2%&&;6B 9
#K
#
-
0
(
# 'CC56#
(
#
'
2
6
.
"
!
,
A/
N
C
N
(
/
9 )
#,
! "3"F
G
%%=
#
,
(
;&
# ,
0
#
L
'C'&#
, - 0)
-
0
,
7
#
$
/
6 $
(
#
-
,
0
2
(
! "3#5
(
0
7
"
)
0
0,
6# /
2
#
,
# 'C4>6
#
2
0
0
#
/
"
1
,
K
/
2'C><6 1
.
(
(
0
,
7
(
(
Pseudolagosuchus major, Arcucci, 1987. Vista
lateral del material holotipo, con fragmentos de
cuerpos vertebrales dorsales, pubis, fémur, tibia,
fíbula y vértebras caudales proximales. Tomado
de Arcucci.
#
!
/
#7
0
@ /
(
#
(
7
2'CC56#
@
2! %F"F
G A! %FF9
(
%&&=6#
%#= #
,
0
#
(
#
(
/ 0
%=&
/
#
)
H
(
/.
#
,
2
0
0
(
- #
,
L#
.
G
9 )
(
9
/
K
(
,
/
#
#
! %FFB
2N
(
3
#
1 #
#
(
E(
2$
0"
# %&&;6#
)#
0
0# '4446#
/
(
#
#
,
#0,
,
,
0
,
/
1
#0,
7
/
,
(
)
(
@
@
9 )
#
.
K ,
0=
@
>
.
.
(
#0(
(
,
%%< %'&
2
#
#
(
,
K0
$
E/
)
0 @
6(
'C45
"
08
! "338
,
//
#
(
,
0
,
%
0
'CC<
,
/
(
)
!
7
/
/
#
(
N
,
N
0 /
0
"
(
'<
0
,
-
2
,
)
.
8
@
%=<
,
-
0
)6
)(
0
()
( ' ( )
(
,
0 /
7 ()
0
/
#
#
,
#
#
9 )
,
#.
/ (
9
$
/
9 )
8
#
5
#
(
- 0
)
0,
- 0
,
#
0,
)
/
1
7
! "3B9
Gentileza de Daniel Banchero
'C;& 0
/.
,
# 0
#
#
'C;=
)
!"
"
# $ "
) + 0 ( -
.
%!"
"
# $ "
/
"
G
%
(
/.
#
!
0
$ 2 /
#
$ #
(
,
7
6#
(
#
#0
)
G
/
G
0
@
(
)
#"
#
#
)
(
,
#
(
Z
"
.
,
7
/
#
/ #
1
)
"
"
7 #
,
G
#
,
1
7
0
(
1
#
,
!A 0 A
0(
: )
7
Los dedos centrales de la mano de Herrerasaurus terminan en grandes garras
marcadamente curvas, semejantes a las que las aves rapaces presentan en sus
pies. Las garras eran flexionadas por músculos que se insertaban en
prominentes tubérculos proximoventrales. Los dedos externos, atrofiados, eran
inútiles en la manipulación. Cada uno de estos detalles anatómicos concuerda
con los que exhiben otros terópodos primitivos, como Coelophysis, a la par
que diferencian a Herrerasaurus de ornitisquios y saurisquios
sauropodomorfos, en los cuales las manos son más cortas y la atrofia de los
dedos externos no es tan marcada.
2 2! "339
Gentileza de Daniel Banchero
.
,
1
//
9 )
'
G
2%=& 0 %%<
0 =&
)
#
/
$
1
/
6#
,
,
- 6#
#
8
#
CL
/
5
#
2
#
,
"
0
,
0
(
0
/
1) @
! "35F :
Gentileza de Daniel Banchero
9 )
7
2%%< %='
$
,
(
#
- 6
N
$
)
8
M
,
$
(
K
0
&
2K
# 2%&&>6
2K
&
,
"
#K
,
M
0
7 ,
(
,
,
#
2
/
,
#K
-
0
N
# 'CC46#
(
#&
2! "33#
7
(
$
6
)
(
,
6
0
,
#
//
M
%%> %%&
$ #K
@
#
#
'
%'5
0
9
"
(
.
#
/.
0
29 )
$
9
#
0
,
.
/
#@
7
2! %FF3
6# 0 ,
0
9 )
,
#
%%4
#
.
G #
0
)
@ / 8.1
7
.
2@
.
6#
,
$
/
28
@
, . #
6# /
Q
1
#
,
#
G
/
GE
%&&C6#
(
#
)
,
9
@ ! %FF3
$
%%4
G
/
(
H
2/
/
29 )
# 'C=%6
$
#
/
7
)
/
(
(
0$
0
#
7
#
#
'
G
# 'CCC6#
$
$
-
2! "333
#
,
(
,
(
(
//
N
'#<&
#
%%=
$
8
<&
-
2
M
0
6 $
$ #K
2"
.
'& L
9
(
/
"
/
/
2K6 3
#
(
(
/
2%6
#
(
2 6 $L
L 2(
(
256
0 ((
(
246
2(
M
# 'C4< 6
0
2 6
]
# 9]
2V
/
# %&&= 6#
2'6 (
(
(
2<6
(
# 2C6
(
2>6
#
(
I
(
QV
L
# 'C4;6# 1
(
#
2'&6
2''6
1
0(
2M !9 !
!!!6 (
B K ^ $8@$ '%C5C $
(
L
(
#
2N6
2 6"
/
(
2 6
2;6
#
2=6
/
0
2M6
(
1
/
(
[[$
B
\\ !
^' PK #M
( (
(
2 6
#
(
/
(]
^<
M
P N ^ '&
(
)
#
! "3B5
)
/
1
>I
0
$
#
/
/
*
,
"
7
.
0(
(
/
9 )
6#
+
(
,
,
(
2
/ 0
)
,
(
0
(
1
"
1
,
#
/
)
,
#
/
)
/
2
,
N
!
,
0
#0,
/
# (
"
,
/
$
,
6
#
(
)
#
7
0
(
9
,
/
/
2
)
,
/
#
K
,
$
(
!
N
/
2N6#
(
/
1
(
0
(
P 2 6#
7
(
2 6#
/
(
P 2M6#
/
/
2 A" %<>>6 (
H
9
(
1 P 2K6#
/
(
1 P 2 6#
7 P 2 6#
/
( (
7 P
/ P$
, '
/
/
/
$
0
(
(
0 (
^'
#K
2
2 #
(
\ 2'C>;6
6P K# $
\ 2'CC'6
/
B (# (
P #
(
K
# 'C>;P K#
(
'CC' 6
P #
(
$
P #
#
$
(
!
N
(
/
/
(
0
(
(
2 6#
P 2:6#
P 2"6#
1
2 A" %<>>6 (
/
(
/
(
/
(
$
H
9
P 2!6#
P 2W6#
, '
(
#
/
/
/
P #(
P #
(
/
(
/
2N
P K#
/
P N#
/
K
$
^'
# 'C>; 6
P #
,
! "35#
(
0
! >
!E
>
! %FF5
( #
# K# "
#
( /
.(
R
(
(
/
6
,
)
G
0
,
$
03
$
. (
,
#
,
#
,
(
.
//
R 1.2
#0
(
#
( .
#
2
0
/
,
/
! "33$2
G
)
4 %"5! 67 2
89:
,
( .
#
.
#
/ #,
#
0
) + 0 ( -
(
I
6
J
?
?K>
I
@
G
6
I
6
6
>
.
7@
@H6
>7
@
L
6
>
2
6 H?
>
>
!
2
>
?
J
>
6
>7 @
>6 G
6
-
"
)
+
/
'
+ 0
)
/
#
/
1
/.
0
)
/.
/.
1.2
! "33$2
P
/
.
)
/
#
,
/
4 %"5! 67 2 8$:
( ) ( /
"
'
+ 0
/
(
,
= < /.
(
/
0
(
/ ,
G
,
#
(
0
(
,
M7
/ 0
1
,
(
((
1
(
#
- - 0)
1
#
,
G
/
(.
0
)
(
(
, /
+
0
(- 0
#
#
/
(.
G
0
#
7
-
)
.
0
'
( (
/
(
2#
$ 6
G
$ #
#
(
(
" #
7
$
G
G
/
2%
#
#
G
)
$ 6
7
#
#
)
%
0
#
,
/
,
/
9
.
(
0
(
/
)
0
/
,
(
( /
/ ,
/
.
,
,
-
"
(
(
) ,
)
)
+
1
0
+
#
, (
/
2
,
)
#
)
(
(
0 )
6#
)
,
,
)
#
#
#
,
)
0 )
,
)
0
.
/
0+
, (
( N
En Dinosauria el extremo distal del isquión forma una barra de sección
subtriangular, a diferencia de la mayoría de los reptiles, ornitisquios
incluidos, en los cuales es transversalmente aplanado.
)
,
#
)C
(
7
)
N
)
)
0
/
N
(
G
%=5==
/
+
,
#
+C
7
0
1
N.
)
(
)
/
(
)
,
1
0
#
7
N
9
0
9
"
á
0
#
2
#
0
7
,
6
/
/
,
1.2
í
ó
6á 2 8$:
0
! "33$2
0
)
í
4 %"5!
/
,
/
Ní
-C
.
#
(
7
)
/
.
,
.
(
/ #
/
(
7
)
(
)(
(
)
_
(
/
/
,
#
(
2
)
!
) /.
(
1
K
G
G
, )
(
(
0
/
/
/
G
!,
(
)
/
1
G
2
6
8
#
H /
Centro de Estudios Paleontológicos de Chile
Roberto Díaz Aros
Centro de Estudios Paleontológicos de Chile
[email protected]
www.cepchile-paleontologia.es.tl
0 0( ,
! 2
&" *
0 AA ! 2+2
.
<
.
)
"3#5 2 R
"
29
6(
"
N
2
#8
9
6# "
7 #
R
* +, $
- ./ +01.-2 3414/! 5
! 0436
,
! -2 "35$ 2 *
00
(/
+
*
! 7 ./32083106.
0
! /2 O2 %FFB 2 *9
(
+
+%
$
(
9
1
9
! -2
001
04
#
!5
!
:
0
6
! O2 .2! "358 *@ /
"
0
+
& 9
! $ 0;2014<
) >I
2/2 "3B5 2 *H
/
,
)
2
#
3
6
N
!
#
+
! =+.-2/618/
G A! O2 %FF9 2 *
L
/
((
(
0
"
9
(
+
#7
# %92=6B <<; <>5
G
! /2 2 %FFB 2 R
/
(
0
(
(
0
$ / Q"
# 'CCC (
H
9
(@
8 1 # H$ R & ) 7 ,
.<<8 .3 +/- >
#(
?
@"
!
.
! O2P E (
! /2)2 %FFB R "
9
(
(
K
(
0
R "
5
'C 2'6B 'U''
,
! O2 2 "3#B 2 *$
00
(
+ 2
# W 2 6B
)
#5
$
#A
#
% #
#
#B 'U<<
> ! 2 02P - A ! 2 ,2P
@ ! 2 O2 E ( ! O2 %FF5 2 R
0
B
9
R "
5
! .<<6B 04+0-2 ;1..
(
! /2)2! @
! .2!
! /2!
0
! /2 "333 R
(
H
9
2
6 (
K
R%
,
C
?
! )*+2 =001B=06
/ H G! 2 2 E
@ ! 2 2 %FF3 * K
$
2
B$
6
(
!
N
29
#
6
0 /
(
$
+
) ! $ /!
. /;46
@ ! 2 O2P > ! 2 02P - A ! 2 ,2P > ! 2 2P +
! 2 C2 E Q ! +2 %FF3 2
R
0
(
(
"
9
(
@
R
#7
%3 2%6B 5C4U<';
/( )+
@ ! 2 O2P > ! 2 2P > ! 2 02P +
! 2 C2P Q
2E
/2 2 %FF3 2 R
$L
("
9
$
0 /
(
+
!
$ ;.8 00 0. .<<4
! .2 2 "33% 2 * 0
(
#
+
! 7 ;=2=018.
.2 2 "33$ 2 *3
! .2 2 "33$@ 2 *@
+
(
2H
9
! 2/2 2 E 2 C
!
6 (
! 2 2 "3B9 2 *"
+ 28
)
6
6+
2
N
> ! 2 2 "35% 2 *9
(
! -2 2 "335 2 *9
B
%
! *D .062 =.1=4
L
(
!
! 7
6(
+
0;2/<<1/.;
2 %FFF R
/
M # "8B >< 44
@S
> ! 2 2 "35" 2 *9
9
0
29
N
$
-
/
#7
(
,
$ :
! ;! 2;1.<
69
+ 5 $
2
9
*
0"
N
! ;662013
69
/
(
0446 .=2/;=F34
R
7 2
!J 9
N
JA N
E 5 $
;4/2016
+
, $
G
-
Descargar