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ra. Laura Böhm Stoffel - Ing. Agr. - Peter Seemann Fahrenkrog - Ing. Agr., Dr .rer. hort. & " ' ( ) 9 ##+# ! "" ## $ " % "& " D, '! * & " ' ( $# $# El autor desea expresar sus más sinceros agradecimientos: Al Ing. Agr. M.Sc. Ph.D. Sr. Luigi Ciampi Panno por su abnegada participación y orientación durante la presente investigación. A BOPAR S.A por facilitar el material biológico necesario para desarrollar la investigación. A Karin Hoffens y Rodrigo Mardones por facilitar el material biológico necesario para desarrollar la investigación. Al personal del laboratorio de Fitopatología del Instituto de Producción y Sanidad Vegetal, el cual siempre prestó valiosa colaboración durante el trabajo práctico. A todas las personas que de una u otra forma hicieron posible la realización de esta tesis. & " ' ( + 9 ##+# ! "" ## $ " % "& " D, '! , & " ' ( 1' 22 1' 22 - ./ $ 0" 0 " (" 1 2 3 ( 3 1 3 " 3 4 5 3 3 0 3 / 6 6" (7 - ./ 8 2 6 8 $ 72 3 3 ( 3 3 $" $ % 6 3 $" $ (" 9 ( : ;( " ( (" 7 < 3 8 ( 3 =3 > " / 2 3 8 ;(" 9 / ( : ( 3 " $ 8 - ./" $ 0 ( ( $ 3 ( ( 3 & " ' ( ? 9 ##+# ! 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itopatología. México. Limusa 756 p. ARK, P. y M. STARR. 1951. Bacterial diseases of orchids. Plant Disease Reporter 35: 42 - 43. BATEMAN, H. y R. MILLAR. 1966. Pectic enzymes in tissue degradation. Annual Review of Phytopathology 4: 199 - 146. BESOAIN. X, M. SAAVEDRA, G. VERDUGO, E. BRICEÑO, L. BAEZA y G.CHAIN. 2002. Pudrición blanda cremosa de la cala de color (Zantedeschia sp.). In: Libro de Resumenes XII Congreso Sociedad Chilena de Fitopatología. Puerto Varas 8 - 7 oct. p 78. BIGRE, J.P, MORAND, J.C y THARAUD M.1990. Patología de los cultivos florales y ornamentales. Traducido por Antonio Peña - Iglesias. Madrid, España. Mundi Prensa. 233 p. BLOM, T.J y BROWN, W. 1999. Preplant Copper - based Compounds Reduce Erwinia Soft Rot on Calla Lilys. Hor technology 9 (1):51 - 59. BUDDENHAGEN, W.1970.The Relation of Plant Bacteria in the Soil.In: Baker, F. y Snyder, W (eds). Ecology of soil - borne Plant Pathogens. Prelude to Biological Control. University of California, Berkeley, Estados Unidos. 571 p. BURR, T. y M. SCHROTH y T. SUSLOW. 1978. Increased potato yields by treatment of seed pieces with specific strains of Pseudomonas fluorescens y P. putida. & " ' ( AI 9 ##+# ! "" ## $ " % "& " D, '! Phytopathology 68: 1377 - 1383. BREWER, J., M. HARRISON y J. WINSTON. 1980. Differential attraction of Drosophila melanogaster Meig to potato tissue infected with two varieties of Erwinia carotovora. American Potato Journal 57(5): 219 -224. CIAMPI, L. 1972. Enfermedades bacterianas de la papa en Chile. Ecología y distribución. Fitopatología 7: 5 -14. -------- y G. PASSALACQUA. 1983. Bacterias del género Erwinia (grupo carotovora) aisladas de plantas y tubérculos de papa (Solanum tuberosum L.). Phyton 43(2):221 226. -------- .1990. Enfermedades bacterianas del cultivo de la papa: Aspectos etiológicos, epidemiológicos y control. In: IV Curso Internacional de Producción y Almacenamiento de Papa. 26 de noviembre al 14 de diciembre. INIA - CIP - PNUD, Osorno, Chile. s/p. ---------.2002. Universidad Austral de Chile. Introducción a la Patología Vegetal. Valdivia, Chile. Nuova Firenze.232 p. DAVIS, R y J. NUNEZ. 2003. Potato Bacterial Soft Rot and Blackleg. University of California. <www.ipm.ucdavis.edu/PMG/r607100111.html. (20.oct.2003). DE BOER, S. y A. KELMAN. 1975. Evaluation of procedures for detection of pectolytic Erwinia spp. on potato tubers. Potato Research 52: 117 - 123. DE BOER, S. y A. KELMAN.1978. Influence of oxigen concentration and storage factors on susceptibility of potato tubers to bacterial “soft rot” (Erwinia carotovora). Potato Research 21: 65 - 80. DE HERTOG W, y LE NARD, W.1993.The Physiology of Flower Bulbs. Amsterdam, Holanda. Elsevier. 811 p. DOLE, M.J y WILKINS, H. F. 1999. Floriculture, principles and species. New Jersey, USA. Prentice Hall .613 p. DOUDOROFF, M. y PALLERONI, N. 1974. The genus Pseudomonas. In: R. Buchanan and N. E. Gibbons (eds). Bergey’ s Manual of Determinative Bacteriology 8th. Edn. Williams and Wilkins, Baltimore. pp 217 - 243. DYE, D. 1969. A taxonomic study of the genus Erwinia; II the carotovora group. New Zealand Journal of Science 12: 81 - 97. ELPHINSTONE, J. 1988. Methods of control of Erwinia diseases of the potato. In (eds). Bacterial diseases of the potato. Report of the Planning Conference on bacterial diseases of the potato. International Potato Center. Lima, Perú pp: 187 - 192. FAHY, P.C y PERSLEY. 1983. Plant Bacterial Diseases. A Diagnostic Guide. Sydney, Australia. Academic Press. 393 p. GIBSON, T. y GORDON, R. 1974. The genus Bacillus. In: Bergey`s Manual of determinative Bacteriology. R.E. Buchanan and N.E. Gibbons (Eds). 8th . Edn. Williams and Wilkins, Baltimore. pp: 219 - 257. GOTTSCHALK, G.1979. Bacterial metabolism. New York. Springer Verlag. 281 p. GRAHAM, D. y W. DOWSON. 1960. The coliform bacteria associated with potato blackleg and other soft rot I. Their pathogenicity in relation to temperature. Annals of Applied Biology 48: 51 - 57. AC & " ' ( =' 33245 -------. y HODGKISS. 1967. Identity of Gram - negative, yellow pigmented, fermentative bacteria isolated from plants and animals. Journal of Applied Bacteriology 30: 175 189. HALL, J. y R. WOOD. 1970. Plant cells killed by soft rot parasites. Nature 227 (19): 1266 - 1267. JACOBS, F.1997.Calla en maceta y calla para flor cortada, dos nuevas posibilidades para el mercado español. < http: // /calla.html (5 abri. 2003). KLEMENT, Z. RUDOLPH, K y SANDS, D. C. 1990. Methods in Phytobacteriology. Budapest, Hungria. Akademiai Kiado. 568p. KORN - WENDISCH, F. y H. JÜRGEN. 1992. The family Streptomicetae. The Prokaryotes. A Handbook on the biology of bacteria: Ecophysiology, Isolation, Identification, Applications, 2ed Volumen I. New York, Springer - Verlag. pp: 921 995. LIAO, H. 1989. Antagonism of Pseudomona putida strain PP - 22 to phytopathogenic bacteria and its potential use as biocontrol agent. Plant Disease 73: 223 - 226. LIPTON, W. 1967. Some effects of low - oxygen atmospheres on potato tubers. American Potato Journal 44: 292 - 299. LUND, B. y J. NICHOLLS. 1970. Factors influencing the soft rotting of potato tubers by bacteria. Potato Research 13: 210 - 214. -------. y G. WYATT. 1972. The effect of oxigen and carbon dioxide concentration on bacterial soft rot of potatoes I. King Edward potatoes inoculated with Erwinia carotovora subsp. atroseptica. Potato Research 15: 174 - 179. MOLINA, J., M. HARRISON y J. BREWER. 1974. Transmission of Erwinia carotovora subsp. atroseptica by Drosophila melanogaster Meig I Acquisition and transmission of the bacterium. American Potato Journal 51: 245 - 250. MOLINA, J. y M. HARRISON. 1980. The role of Erwinia carotovora in the epidemiology of potato blackleg. II The effect of soil temperature on diseases severity. American Potato Journal 57: 351 - 363. MOUNT M.S y LACY, G. H.1982.Phytopathogenic prokaryotes. New York, USA. Academic Press.541 p. PEROMBELON, M.C.M y LOWE. 1975. Studies on the initiaton of bacterial soft rot in potato tuber. Potato Research 18:64 - 82. --------.y A. KELMAN. 1980. Ecology of soft rot Erwinias. Annual Review of Phytopathology 18: 361 - 387. --------. 1987. Pathogenesis by pectolytic Erwinias. In: Civerolo. E, Collmer. A, Davis. R y Gillaspie. A (eds). Plant pathogenic bacteria. Proceeding of the sixth International Conference on Plant pathogenic bacteria, June 2 - 7 1985. Netherland, Nijhoff Publishers. pp: 109 - 120. --------. 1988. Ecology of Erwinias causing stem and tuber disease. In: (eds). Bacterial disease of potato. Report of the Planning Conference on Bacterial Disease of the potato. International Potato Center. Lima, Perú. pp: 143 - 177. -------. y VAN DER WOLF J. M.1998. Methods for the detection and quantification of Erwinia carotovora subsp. atroseptica on potatoes. Scotland. Scottish Crop Research & " ' ( AB 9 ##+# ! "" ## $ " % "& " D, '! Institute. 76 p. PIRHONEN, M. 1992. Characterization of pathogenicity determinants of the plant pathogen Erwinia carotovora. Uppsala.Swedish University of Agricultural Sciences. 78 p. --------. D. FLEGO, R. HEIKINHEIMO y E. PALVA. 1993. A small diffusible signal molecule is responsable for the global control of virulence and exoenzime production in the plant pathogen Erwinia carotovora. The EMBO Journal 12(6): 2467 - 2476. PIZANO, M. 1999. Zantedeschia, Calla Lily. Santafé de Bogotá, Colombia. Horticiencia. 54 p. RHODES, D. y LOGAN. 1987. A method for selecting fluorescent Pseudomonads inhibitory to seed tuber decay. Potato Research 30: 603 - 611. RIED, J. y A. COLLMER. 1986. Comparison of pectic enzymes produced by Erwinia chrysantemi, Erwinia carotovora subsp. carotovora, and Erwinia carotovora subsp. atroseptica. Applied and Enviromental Microbiology 52(2): 305 - 310. SALINGER, J.1994. Comercial Flower Growing. Inkata. Sydney, Australia. 269 p. SCHAAD, N.W.1994. Laboratory Guide for identification of Plant Pathogenic Bacteria. 2ed. The American Phytopathological Society. Minnesota, USA. 164 p. SMITH, I.M. DUNEZ, J. LELLIOTT, R. A. PHILLIPS, D. H y ARCHER, R. A. 1992. Manual de Enfermedades de las plantas. Traducido por Fernando García Arenal. Bilbao, España. Mundi - Prensa.671 p. SMITH, C. y BARTZ. 1990. Variation in the pathogenicity and aggressiveness of strains of Erwinia carotovora subsp. carotovora isolated from different hosts. Plant Diseases 74: 505 - 509. SMITH, L. y HOBBS, G. 1974. The genus Clostridium. Pp 551 - 572. In: R. E. Buchanan and N.E. Gibbons (Eds). Bergey`s Manual of determinative Bacteriology 8th. Edn. Williams and Wilkins, Baltimore. 343 p. SNIJDER, C. VAN TUYL y JAAP. M. 2002. Evaluation of test to determine resistance of Zantedeschia spp. (Araceae) to soft rot caused by Erwinia carotovora subsp. carotovora. European Journal of Plant Pathology. 108: 565 - 571. STARR, M. y A. CHATTERJEE. 1972. The genus Erwinia: Enterobacteria pathogenic to plants and animals. Annual Review of Microbiology 26: 389 - 426. STANIER, R, M. DAUDEROF y E. ADELBERG. 1965. El Mundo de Los Microbios. Aguilar, España.724 p. TOPLESS. S. 2000. Horticulture news. Nueva Zelanda. 6 p. VANNESTE, J.L, PERRY J.J, PERRY L.J-MEYER y BEDFORD R.J.1994. Erwinia herbicola Eh 252 as a Biological Control Agent of Bacterial Soft Rot on Potatoes. Hort Research, Ruakura Research Centre, Hamilton.<www.hortnet.co.nz/publications/nzpps/proceedings/94/94_198.htm.> (30 abr. 2003). VANNESTE, J. L y D. A. CORNISH. 1995. Development of Quantitative Bioassay To Study The Interaction Between Biological Control Agents And Soft Rot Erwinia. Hort Research, Ruakura Research Centre, Hamilton<www.hornet.co.nz/publications/nzpps/procedings/95/95.> (24. oct. 2003). IE & " ' ( =' 33245 VISSER, R., W. HOLZAPFEL, J. BEZVIDENHOUT y J. KOTZE. 1986. Antagonism of lactic acid bacteria agains phytopathogenic bacteria. Applied and Enviromental Microbiology 52(3): 552 - 555. WEBER, J. y C. WEGENER. 1986. Virulence and enziyme production of Erwinia carotovora subsp. atroseptica on potato tuber tissue. Journal of Phytopathology 117: 97 - 106. WEI CHEN C. Study on the Control of Calla Lily Bacterial Soft Rot Disease. <www.tdais.gov.tw/search/issn/61 - 70/bow69 - 4.htm.> (30. abri. 2003). WELSH, T.1991.The New Zealand Cala. Combined Procedings International Plant Propagators Society. 41: 478 - 484. WRIGHT, P.J.1994. Controlling soft rots in callas. < www.crop.cri.nz/psp/articles/docs/callarot.htm. > (26 abri. 2003). -------. y BURGE, G.K.2000. Irrigation, sawdust mulch, and enhance biocide affects soft rot incidence, and flower and tuber production of calla. New Zealand Journal of Crop and Horticultural Science.28:225 - 231. --------. y TRIGGS C.M, 2002. Effects of cessation of irrigation and time of lifting of tubers on bacterial soft rot of calla (Zantedeschia spp.) tubers. New Zealand Journal of Crop and Horticultural Science. 30: 265 - 272. & " ' ( I)