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DEDICATORIA
A mis queridsimos padres, Wilfredo y Yolanda, por su infinito amor y
comprensin, a mi hermano Wilfredo y a todas aquellas personas por su apoyo incondicional,
especialmente a Hamilton. A los cautivantes bosques nublados por su enigmtico encanto que
insta a conocerlos ms y a respetarlos.
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Este trabajo es posible gracias a la oportunidad dada por el Mg. Asunción Cano y el Dr. Víctor
Pacheco, quienes confiaron en mí para la realización de esta tesis. Debo agradecer en especial al
Mg. Asunción Cano por acogerme en su laboratorio, orientarme y brindarme su apoyo durante el
desarrollo de la tesis.
Al Dr. Víctor Pacheco, investigador del Museo de Historia Natural de la Universidad
Nacional Mayor de San Marcos, por hacerme partícipe del proyecto “Protección de la
Biodiversidad en Bosques Montanos Fragmentados y Propuesta para Conservar el Bosque de
Carpish, Huánuco”, ganador del Concurso Nacional por Proyecto de Investigación, “Santiago
Antunez de Mayolo – 2001”, auspiciado por el Consejo Nacional de Ciencia y Tecnología
(CONCYTEC) y mediante el cual se pudo financiar parcialmente la tesis.
Agradezco de manera especial al Mg. Hamilton Beltrán del Herbario San Marcos (USM),
por su gran e invalorable ayuda en el trabajo de campo y orientación durante el desarrollo de la
tesis.
Así mismo agradezco al Dr. John Clark y al Dr. Laurence Skog del Smithsonian Institution,
Dr. James Smith del Boise State University y a la Dra. Blanca León del University of Texas in
Austin, quienes me brindaron bibliografía especializada y ayudaron en la confirmación de las
determinaciones, en particular al Dr. John Clark por ser mi maestro y afianzar mi interés en las
gesneriáceas.
Al Mg. Rodolfo Vásquez y a la Blga. Rocío Rojas del Jardín Botánico de Missouri por
darme la oportunidad de poder iniciar mi investigación en botánica en los bosques nublados.
A Marina Villalobos, Miguel Chocce y todos los compañeros de las expediciones por su
amistad y hacer de las salidas de campo una experiencia gratificante. A Jorge Lingán, Marybel
Morales y Maritza Rodríguez por ser estímulos constantes de superación y compartir su amistad.
Agradezco a la Dra. Magda Chanco, a la Mg. María I. La Torre, a la Mg. Joaquina Albán, a
mis compañeros de laboratorio, José Roque, Ángel Ramírez y Wilfredo Mendoza, y a Arturo
Granda por sus valiosos comentarios y sugerencias.
Doy un agradecimiento especial a los pobladores de la Comunidad de San Pedro de Carpish
por su hospitalidad, confianza y prestancia para ayudar durante las expediciones científicas.
Un infinito agradecimiento a mis padres por apoyarme a conseguir mis metas, es de ellos
también esta tesis, gracias Wilfredo y Yolanda.
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Se presenta el estudio taxonómico del Orden Scrophulariales en los bosques montanos húmedos
de Carpish, departamento de Huánuco. Se realizó colectas entre los 1700 a 3100 m de altitud
bajo técnicas estándares de estudios florísticos. El estudio taxonómico consistió en la elaboración
de claves para familias, géneros y especies, y en la descripción, análisis taxonómico y
distribución geográfica para cada especie. Se reportan 4 familias, 20 géneros y 38 especies; la
familia más diversa es Gesneriaceae con 21 especies en 8 géneros, seguida de Scrophulariaceae
con 12 especies en 8 géneros; Acanthaceae con 3 especies en 3 géneros y Lentibulariaceae con 2
especies en 1 género. Se han dado cambios nomenclaturales en algunas especies, con excepción
en Lentibulariaceae. El género más diverso es Alloplectus, de la familia Gesneriaceae, con 8
especies. Las especies A. carpishensis, A. ichthyoderma y Utricularia unifolia, esta última de la
familia Lentibulariaceae, tienen mayor rango altitudinal (hasta los 3000 m). A. carpishensis es
una especie nueva para la ciencia. De las especies estudiadas en los bosques de Carpish, 9 son
endémicas del Perú, la mayoría (5) pertenecen a la familia Gesneriaceae. Las especies A.
dodsonii y Columnea fuscihirta, de la misma familia, son registros nuevos para la flora peruana.
Diez especies amplían su rango de distribución al departamento de Huánuco.
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A taxonomic study of the Scrophulariales is presented in the humid montane forests of Carpish,
Huanuco department. The collects were realized between 1700 and 3100 m, following standard
techniques of floristic studies. The taxonomic study consisted in the elaboration of keys for
families, genera and species, and description, taxonomic analysis and geographic distribution for
each one of the species. Four families, 20 genus and 38 species are reported; the most diverse
family is Gesneriaceae with 21 species in 8 genus, followed by Scrophulariaceae with 12 species
in 8 genus; Acanthaceae with 3 species in 3 genus and Lentibulariaceae with 2 species in 1
genus. Nomenclatural changes have been given in some species, with exception in the family
Lentibulariaceae. The most diverse genus is Alloplectus, of the family Gesneriaceae, with 8
species. The species A. carpishensis, A. ichthyoderma and Utricularia unifolia, this last of the
family Lentibulariaceae, can inhabit to more altitudes (until 3000 m). A. carpishensis is a new
species to the science. Of the species studied in the forests of Carpish, 9 are endemic of the Peru,
the majority (5) belong to the family Gesneriaceae. The species A. dodsonii and Columnea
fuscihirta, of the same family, are new registrations for the Peruvian flora. Ten species enlarge
their distribution range to the department of Huanuco.
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